Lancia ran cars in Safari, Acropolis with a pre-filtering system in the boot of the car. Early setups ran the car with the boot-lid jacked up to let air in. Later cars had moulded-in ducts from the vents under the roof spoiler back into the boot. Flexible ducts then connected through the bulkhead from the pre-filter to either side of the air-box.
Possible that that is what these are for (to let air in), though i did see in one of the photos that the air-box is not an enclosed one so no way to connect a pre-filtering system. Easy to change over when needed though. Other than that, no idea really !
David